Aspect Ratio.
There is plenty on it.
No I am not fired up, its simply that the argument that a "better" photo is made with a smaller sensor is at least to my experience, a non issue.
When you are dealing with the high levels of cropping that occur in most people's wildlife photographs, you can bet that it is an issue. One can increase resolution or buy a bigger lens. Increasing the lens size is ideal—but: the step size from 400mm to 800mm might be from $1,000 to $20,000. Plus the lenses weigh a ton.
Again, I think you have no idea what the issues are in wildlife photography.
OK...
Look.
35mm is 1.5-1.6 times larger than APS. (Nikon and Canon respectively)
Both 35mm and APS shoot an aspect ration of 3:2.
There is distortion in all lenses that the further to the edge you get in any lens the more distorted things become.
The further from the center you go there is distortion and a form of "skewing" where things elongate from the lens aspects. This simply cannot be eliminated. It can be minimized.
When you are trying to compose an image of say a bird in flight, and your shooting 35mm, while framing the image as close to center as possible, there is going to be an amount outside the center that will regardless of equipment used, get the skewing going on.
So if you frame the image in the FF sensor to the equivalent of an APS sensor, the image in the 35 (FF) image is less distorted and ergo when the final print comes out is closer in actual proportions than the APS.
In an APS sensor, that same image is more greatly distorted. Ergo, the wings if compared (if possible) would not look quite the same as in the 35mm regardless of the cropping.
The pixle size becomes irrelevant because the distortion factor in relation to sensor (format size) takes the game.
When I talk about projection, it is a factor found in cartography.
The further from the center you get, the more distortion takes place because in projection, your representing a 3D object in 2 dimensions. This same effect is occurring in photography because your trying to capture a 3D object into a 2D plain.
So distortion will exist regardless. One simply does not see that distortion unless one were to shoot the exact same object with diff. format sizes.