LR2 does not. It uses MelissaRGB a modified version of ProPhotoRGB that is entirely linearised just like the data in your RAW file. The theory is the initial conversion to MelissaRGB allows their controls to work to the best extent where things like highlight Recovery depend a lot on the ability to mathematically edit more colours that monitors can display or printers can possibly print, and more importantly it does all this at the maximum bitrate of the RAW file.
Only when you export does it pick a profile for you. Ultimately the idea is if you want maximum colour range reproduction then you pick a profile that is larger than the colour range of your image. In 99% of cases my images fit nicely into sRGB so even AdobeRGB is overkill. The colour gamut of standard matte paper also is just barely larger than sRGB so unless you're using some awesome printing process and a pro lab, or have a nice wide gamut monitor there's no reason really to battle with the larger gamuts.
That's not really the issue here though. Colours which fall outside the gamut go through a conversion process. For photography this is called the "Perceptual" rendering intent. The colours are clipped at their maximum, and the most saturated colours around the image are scaled ever so slightly so that the result does not look like clipping. The end result is entirely artefact free, looks natural, and covers the target gamut nicely.
As a matter of interest are you working with an original RAW file, and how much editing have you done to the image? There are a few factors that could be in play. Posterisation from editing the file too much. Posterisation from editing a file in 8bit rather than in the original or larger bitrate. If the saving is the problem, well most if not all lossy compression algorithms sacrifice the dark and the blue channels since the eye is least sensitive to these. One thing thought, your image does have artefacts, but it's a question of why are you seeing them so much.
I can see them but only just in the original. At the brightest point in there the red channel has a value of 22. I expect this would be plainly noticeable on any well calibrated screen in a dark environment. Also are you viewing your screen dead on? The problem with laptops is the gamut varies with viewing angle very dramatically. For instance my work laptop (admittedly a bad screen) when I view a solid 128 middle grey head on, the top of the screen would be somewhere around 180 and the bottom around 80, it is that bad. So taken note of what it looks like at different viewing angles and share, I haven't used a mac book pro.
Your image with the contrast boosted.